General fitness, health and nutrition · Public discussion

Complexity of aggression

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  1. I'm looking for any resources arguing aggression is not
    evolutionarily adaptive anymore and theoretical proposals of
    what to do about it. I understand there is no "fixer" and
    that is fine but certainly philosophy can address this
    subject. My theory is man, despite his cognitive
    superiority, shares much in common with other animals in
    aggression. It is further my theory that science and
    technology has threatened and indeed is unraveling what "it
    is" to be a human being. Most people have a set of views of
    what "human nature" is and science and technology challenge
    that. Our "human nature" as it is constituted is too simple
    to digest the changes brought about by science and
    technology. We can't assimilate these changes. As a result a
    growing percentage of people become desperate and the
    ultimate reaction is catastrophic.

    Aggression in some ways is adaptive but in other ways it is
    not. Clearly, some aspects of aggression will have to be
    genetically engineered and this will most likely effect many
    other areas, including those areas of aggression which have
    been adaptive. But I don't forsee an organism with its
    gonads in a bucket and a brain on a platform.

    Continuity vs. (Political) Correctness: Animal Models and
    Human Aggression

    D. Caroline Blanchard, Mark Hebert, and Robert J. Blanchard
    E. Blanchard is Research Professor at the Pacific Biomedical
    Research Center and Professor of Genetics and Molecular
    Biology at the John A. Burns School of Medicine of the
    University of Hawaii. Hebert is Assistant Professor of
    Psychology at the University of Hawaii. R. Blanchard is
    Professor of Psychology and Neurosciences, University of
    Hawaii. With HFG support, D. Blanchard and R. Blanchard
    are writing a book on the biology of aggression.

    One of the sadly enduring themes in aggression research is
    the inability of scientists researching humans and animals
    to make meaningful impressions on each other's understanding
    of the behaviors on which they focus. This problem is not
    restricted to aggression research, but it appears to be more
    strongly expressed in this area than in the study of most
    other behavioral phenomena. Researchers on animal or human
    aggression have tended to remain separate from and
    disdainful of each other, even while attending and
    participating in the meetings of a single dedicated society,
    the International Society for Research on Aggression.
    Needless to say, this situation has impeded the development
    of any sort of unified view of aggression that applies to
    both humans and nonhuman animals. It has thus been one of
    several major problems that have hindered progress in
    aggression research during a period in which many other
    somewhat similar areas, for example the neurobiology of
    defense and its relationship to human emotionality, have
    made considerable advances.

    Aggression research does have some distinctive problems in
    bridging the gap between animal and human behavior. One is
    that the concept of aggression as a phenomenon with
    substantial biological underpinnings is widely perceived as
    running counter to many worthy social and political views.
    The assumption underlying the distancing of many of the
    proponents of such views from too close contact with animal
    aggression research is correct: Animal work often suggests
    that aggressive behavior has extensive roots in biology, and
    it is quite likely that such research may raise questions
    about the modulation of human aggression that are difficult
    to answer from the perspective that there is no direct
    biological influence on this behavior.

    These antibiological views encompass a range of concepts,
    from the position that war, specifically, has no biological
    underpinnings, through denial of any direct involvement of
    biology in the substantial interindividual (socioeconomic,
    ethnic, subcultural, gender, age) variation in violence and
    violent crime. But since individual differences in human as
    well as animal aggression are undeniable on a
    phenomenological level, and since these so often seem to be
    associated with factors that clearly relate to biology,
    such as gender and age, the possibility of interactions
    between biology and experience may be admitted. Gender
    differences may be interpreted as reflecting greater
    opportunity for aggression or greater reinforcement of
    aggressive behaviors for boys/men as opposed to
    girls/women. Eiither source, in turn, may derive from
    cultural norms (a relatively "biology-free" explanation) or
    from differences in gender-typical group composition or
    social activities (an account that suggests, but does not
    dwell on, a biological origin for these differences).
    Similarly, while accounts focusing on self-esteem and
    social skills as modulators of aggression may acknowledge
    that these reflect an interaction of personality factors
    such as behavioral inhibition with experience, the
    existence of biological differences directly relating to
    aggression is often denied, minimized, or ignored.

    A second problem in acknowledging a relationship between
    human and animal aggression involves the cognitive distance
    between humans and other animals. There appears to be a
    relatively well-developed consensus that no known nonhuman
    species has cognitive or linguistic capabilities that are
    close to those of humans. The difference is highly relevant
    to the study of aggression because many instances of human
    aggression are clearly accompanied by complex cognitions, or
    expressed in terms of mechanisms that rely on cognitive and
    technological achievements, which may have no direct
    parallels in nonhuman species' behavior. Also, the use of
    technology can ensure that a human act of aggression causes
    an immense amount of damage impossible for animals to
    achieve. Both factors may be involved in some discrepancies
    between human and animal findings. For example, in nonhuman
    mammals, alcohol sometimes increases aggression at low to
    moderate dose levels but almost always reduces it at higher
    doses (see Berry and Smoothy, 1986 for review). The human
    literature on alcohol and aggression provides little
    evidence of such a nonparallel relationship. A strongly alcohol-
    impaired human can inflict damage both verbally and with a
    weapon, whereas an equally impaired animal cannot, no matter
    how high its motivation to attack may be. Perhaps the most
    important consequence of the cognitive and technological gap
    between man and nonhuman animals relates to war. War is
    defined both in terms of aggressive action and in terms of
    the social organization and tactical capabilities of the
    opponent groups involved in this action. If these
    organizational and tactical capabilities are inadequate,
    then the aggressive behavior does not represent "war," no
    matter how focused or how damaging, or how clearly it
    involves groups rather than individuals. Such a
    conceptualization enables a statement that war is a uniquely
    human phenomenon, with no direct parallels in nonhuman
    animals, but it does so only on the basis of
    cognitive/technological differences. Regardless of what
    intellectual or emotional comforts may ensue from treating
    animal and human aggression as fundamentally different and
    unrelated phenomena, unless they genuinely have no
    significant connections, the tactic is scientifically
    counterproductive. It removes the possibility of comparative
    analysis and cross-fertilization of hypotheses between the
    two realms. It deprives human-focused researchers of an
    extensive literature using experimental methodologies to
    investigate aggression phenomena, while those animal
    researchers who ignore the human literature inevitably fail
    to learn of findings that could open up new and fruitful
    areas of investigation using nonhuman animals. Is there any
    way that this segregation can be bridged and common ground
    found for serious consideration of phenomena with which both
    human and animal researchers are deeply involved?

    Varieties of Animal Aggression | While the term
    "aggression" can often be used without misunderstanding in
    ordinary conversation, it has proved to have so many
    different meanings, and to be so difficult to
    conceptualize, that Benjamin (1985) selected it (along with
    "intelligence" and "self-esteem"😉 as a particularly
    egregious example with which to demonstrate the
    difficulties of concept analysis. Analyses from the animal
    literature suggest one important reason that a single, and
    universally accepted, scientific definition of aggression
    has been difficult: Several different phenomena are
    encompassed by the term. Offensive Attack | The distinction
    between "offensive" and "defensive" forms of adult
    aggression has received a good deal of systematic attention
    (e.g. Blanchard and Blanchard, 1977; Blanchard et al.,
    1984), with both of these typically being differentiated
    from play fighting (Pellis, 1988) and from predation.
    Offensive aggression occurs in the context of a resource
    (including territory) or dominance dispute and its
    successful outcome is the termination of the dispute
    through "victory," manifested as flight or defeat-related
    behaviors by the opponent. Dominants (consistent victors)
    gain resources, including access to females (a particularly
    important "resource" from an evolutionary perspective) and
    food (Flannelly and Lore, 1977; Blanchard et al., 1984).
    Offensive attack involves a set of species-typical
    behaviors that enable the aggressive animal to contact
    particular body sites on the opponent where bites or blows
    are delivered. Offensive attack can thus be differentiated
    from other forms of aggression on the basis of the specific
    target sites for attack, as well as the behaviors by which
    these sites are reached.

    Defensive Attack | Defensive attack is seen only when the
    subject is defending its own body, not when it is attacking
    another animal to "defend" a disputed resource. It includes
    a salient threat component not seen in offensive attack,
    with loud vocalizations and display of weapons such as teeth
    or claws. The bites or blows delivered tend to be made on
    different body sites on the opponent than those contacted in
    offensive aggression. The successful outcome of defensive
    aggression is discouragement of the body-threatening
    conspecific or predator and discontinuation of its attack.
    This can occur prior to the defensive attack, as the result
    of defensive threat, or following the delivery of a bite or
    blow, particularly to the sensitive eye/snout sites that are
    the targets of defensive attack. While relatively little
    field research has been done on the effectiveness of
    defensive threat and attack, the strong inhibitory effect of
    fear on predation suggests that defensive threat may serve
    as a considerable deterrent (Pellis et al., 1988).

    Play Fighting | Play fighting is common among the young of
    many mammal

    (Pellis and Pellis, 1991b). The behaviors involved in play
    fighting have considerable structural similarity to those of
    adult attack and defense, in that a species-typical attack
    pattern is used to approach and make contact with a specific
    site on the body of the opponent, while the defender
    utilizes species-typical behaviors to make that body site
    unavailable to the attacker. In addition, across-species
    studies suggest that, as in adult fighting, attack and
    defense in play fighting are motivationally distinct
    behavior patterns (Pellis and Pellis, 1991a). However, the
    transition from juvenile play fights to adult fighting does
    not appear to involve a continuity in individual levels of
    attack tendency from one to the other: Males that show the
    highest attack rates during play fighting tend to become
    subordinates rather than dominants (Pellis and Pellis,
    1992a; Smith et al., 1996). Also, play fighting, at

    adult fighting, as the play fighting attack target in a spar
    between males may correspond to an important contact target
    on the female,

    finding that deprivation of play fighting has more of a
    deleterious

    In addition, play fighting defenses in juvenile females may
    involve some

    advances of males.

    Predation | Phenomena related to predation are also often
    subsumed under the rubric of aggression. In the laboratory,
    many rats kill and eat mice, a behavior that, due in part to
    the similarities of rats and mice, suggests conspecific
    aggression. In fact, predatory (as opposed to conspecific)
    attack can occur in species even more closely related and
    more similar than rats and mice: Grasshopper mice kill and
    eat laboratory mice. However, both the target sites for
    attack and the behaviors typical of the attack pattern are
    different from conspecific attacks by grasshopper mice
    (Pellis and Pellis, 1992b). These findings indicate that
    predatory attack can and should be differentiated from
    conspecific attack, even when the combatants involved are
    closely related animals. In addition to differences in
    stimuli, response patterns, target sites for contact, and
    outcome of these various behavior patterns, recent work on
    the anatomic and neurochemical systems associated with some
    of these strongly suggests that the physiology of these
    systems is also different (e.g. Bandler and Shipley, 1994).
    Most of the work on the neurobiology of "aggressive"
    behaviors has actually involved defensive threat and attack,
    with a relatively substantial literature also on "quiet
    biting attack," which likely corresponds to predation. The
    pharmacology of offensive and defensive aggression appear to
    be different, with the former (Olivier, et al., 1991) but
    not the latter (Blanchard et al., 1985) responding
    dramatically to a class of "serenics" with effects at
    various serotonin receptor subtypes. Motivational variables
    also produce different effects on these behaviors, with fear
    reducing offensive attack (Blanchard et al., 1988) and
    predation (Pellis et al., 1988) but not altering defensive
    attack (Blanchard et al., 1980).

    Varieties of Human Aggression

    Offensive Attack | If the function of offensive aggression
    in animals is to obtain and hold some sequestrable resource,
    how does this relate to more complicated human emotions and
    impulses? We have argued (Blanchard and Blanchard 1984) that
    "anger" is the emotional response to a challenge to some
    resource for which the angry individual has a claim. For
    example, data on violence resulting from "love triangles,"
    (Wilson and Daly, 1992) suggest that these reflect attempts
    to discourage challenges to the perpetrator's relationship
    to and control over the love-object, regardless of whether
    the latter, or a third-party challenger, serves as victim.
    Similarly, Katz's (1988) descriptions of the subjective
    rewards of criminal and violent behavior suggest that the
    "resources" gained by violence often involve status and
    access to particularly valued women and other status symbols
    rather than money or goods. For many habitually violent
    individuals, a challenge successfully overcome is associated
    with strongly positive emotions; the more able the
    challenger, the sweeter the victory. This analysis is
    couched in terms that may seem inappropriate when applied to
    animals, but it might be considered that for violent
    offenders and, for example, male rats responding to an
    intruding challenger, the stimuli and situations eliciting
    aggression, and the consequences of successful and
    unsuccessful aggression, are very similar.

    These considerations also suggest why a "challenge" is so
    frequent and potent a stimulus to elicit human aggression.
    As Daly and Wilson (1988) have pointed out, in populations
    of young men (who notoriously account for a disproportionate
    share of crimes of violence, e.g. Blanchard and Blanchard,
    1983; Campbell, 1995), failures to respond to challenge
    jeopardize the status of the individual in the group and
    limit his ability to command important resources. A simple
    paradigm—"challenge elicits aggressive response"—emerges as
    a common feature of mammals, particularly young postpubertal
    male mammals. The specific challenge involved is typically
    either to the status of the individual within the group, to
    his access to females or other important resources, or to
    both. Higher primates complicate this paradigm in that other
    group members may get involved, either as seconds (Pereira,
    1989; Silk, 1992) or in an attempt to control and defuse the
    situation (Reinhardt et al., 1986). Humans have contributed
    the factor that challenges may be purely verbal (and often
    quite inventive). They have also created a very encompassing
    form of sequestrable resource—money. The paradigm, however,
    remains, and it is a mammalian pattern, not one found only
    in dysfunctional human social groups.

    Defensive Attack | If offensive aggression has some close
    animal-human parallels, what about defensive attack?
    Although defensive attack is easy to define, it is hard to
    observe in people, for both ethical and practical reasons.
    Polarized hypothetical scenarios, however, elicit consistent
    differences for defensive vs. offensive attack situations.
    [censored]-Stinson (reported in Blanchard and Blanchard, 1983)
    asked male and female students to respond to scenarios
    involving either a physical threat (attack by a stranger in
    an isolated spot) or a resource dispute by indicating the
    likelihood of specific actions or feelings. With reference
    to emotional response, fear dominated in the former, and
    anger in the latter. Physical and physiological responses to
    the two situations also differed, with "freezing" and
    becoming "stiff" or showing "nervous breath" describing the
    fear situation, while becoming "hot" or "burning" was
    associated with the resource dispute situation, as were
    "clinching fist," "staring at," and "adrenalin surge."The
    first choice action for the fear situation was to leave as
    soon as possible, followed by (for women) looking around for
    something to hit the attacker

    Neither of these hitting-related choices was among those
    selected as likely in the resource dispute situation,
    although a strong desire to attack the challenger was often
    cited. What is important is not just that these two
    scenarios elicited a variety of strongly differentiated
    physiological responses and subjective feelings, but that
    they were both associated with a perceived tendency to
    either attack, or to want to attack, the opponent.

    Play Fighting | Play fighting obviously occurs in children
    as well as in the young of most other mammal species.
    Prepubertal boys (Boulton, 1993; Honig et al., 1992; Maccoby
    and Jacklin, 1980), like prepubertal rats (Pellis and
    Pellis, 1990), participate more often in fights than do
    comparable females. Also as in other mammals, play fighting
    and serious fighting in children are different, and this
    difference reflects the actions involved as well as factors
    such as facial expression and the apparent intent of the
    participants (Boulton, 1991a). However, the literature does
    contain suggestions of potential differences between the
    play fighting of prepubertal children and that of rats. In
    rats, the animal that proves to be subordinate in adulthood
    initiates most of the play attacks (vide Pellis and Pellis,
    1992a; Smith et al., 1996), whereas play fighting in middle
    school children tends to involve partners that like each
    other and are closely matched in strength, with both weaker
    and stronger children initiating bouts (Boulton, 1991a,b).
    Another difference is that serious fighting is common enough
    in prepubertal children to be of concern to parents
    (Boulton, 1996), whereas serious fighting is seldom observed
    in prepubertal rats (Smith et al., 1996). Some component of
    this difference may reflect specific learning: In a study of
    middle school English children, many of the behaviors (e.g.,
    karate chop, back kick, scissor kick) seen in play fighting
    but not in serious fighting represent actions that are
    likely to have been learned through observation/imitation,
    perhaps of television programs. The latter two did not occur
    in play fighting among Zapotec children, who did, however,
    show some distinctive behaviors of their own, such as burro
    kick and knuckle rap (Fry, 1987), suggesting that the form
    of playfighting in 8-12 year olds already may have been
    greatly altered by culture-differentiated practices.
    Predation | The view that human predation has a biological
    link to that of closely related mammals is supported by
    findings that a variety of primates predate other
    vertebrates, including mammals (e.g. Anderson, 1986; Kudo
    and Mitani, 1985). In addition, primates appear to be among
    the few mammals that also seek out and kill animals of
    species that predate them, or that serve as major
    competitors for prey (Hiraiwa Hasegawa et al., 1986),
    suggesting that primate hunting is by no means limited to
    animals that are to be consumed. Thus the hunting of large
    and dangerous animals not meant for food, a feature of
    virtually every society that has lived in proximity to such
    animals, also has a clear parallel in nonhuman mammalian
    behavior. Animal and Human Aggressions: Parallel
    Neurobehavioral Systems? We are suggesting that, rather than
    try to establish a basic parallel between animal and human
    "aggression," it might be advantageous to look at the
    concept of aggression as consisting of a number of different
    neurobehavioral systems, at least some of which show
    considerable evidence of continuity between nonhuman mammals
    and people.

    This is not to say that everything included under the human
    aggression rubric will have a direct counterpart in other
    mammals. Obviously, the "aggressive" investor or lawyer or
    businessman has no direct correspondent in infrahuman
    mammals, but this may be because the nouns are
    inappropriate, not the adjective (if the noun were
    "politician" the phrase might have an enhanced
    correspondence). All of these designations reflect a common
    theme, of actions and attitudes that seek to expand claims
    to resources, rights, or influence, in a variety of relevant
    arenas. When particular aggression paradigms are
    individually examined there may be either parallels that
    might be overlooked when an undifferentiated "aggression"
    concept is employed or transformations in the organization
    of that specific aggression paradigm as larger-brained
    mammals with more complex social organizations and technical
    capabilities are examined. This offers the possibility of
    real breakthroughs in relating specific human aggression
    phenomena to their nonhuman mammalian parallels.

    What is likely to continue to be a stumbling block is the
    one that has been there all along, that "aggression" is one
    of the most value-laden terms in any language, and probably
    will continue to be. Thus, while aggression can be a "good"
    thing in the context of pursuit of a valued goal, even here
    it carries the baggage of an implication of activity
    encroaching on the rights of others; the tendency is for the
    protagonist to label precisely the same actions as "defense"
    of the desired goal, making the other guy the aggressor. As
    the old saw goes, virtually every country in the world has a
    ministry, bureau, or department of "Defense" while not one
    has a bureau of "Aggression."

  2. << I'm looking for any resources arguing aggression is not
    evolutionarily adaptive anymore and theoretical proposals of
    what to do about it >>

    I've mentioned this before, but it is worth mentioning
    again.

    Karen Horney divided all inner conflicts into 3 groups.

    One was MOVE AGAINST OTHERS, this group she defined clearly
    and in some length. She also devised a method for treating
    the disorder.

    It would answer your concerns IMO.

  3. "Michael Ragland" <[email hidden]> wrote in message
    "]news:[email hidden]...

    Quoted message said:

    I'm looking for any resources arguing aggression is not
    evolutionarily adaptive anymore and theoretical proposals
    of what to do about it.

    As you are deliberately asking the question rather broadly,
    it is probably unfair to ask you to clarify. Nevertheless, I
    have always been a little uncomfortable with your talk about
    "aggression" because it is such a vague concept.

    Just what behaviors are you classifying as aggression?
    Making distinctions is probably useful, because some
    components of aggression are primarily genetic, whereas
    others are mostly cultural. Are you talking about physical
    aggression exclusively, or do you want to include the kinds
    of bad behavior that is exhibited in this news group? Is
    someone who doesn't give a damn about other people's welfare
    aggressive, or does he also need to be exploitive? To what
    extent can aggressiveness be separated from a belief that
    the world is a competitive place?

    It also might help to clarify if you were to describe more
    clearly what is intended by the "anymore" in "aggression is
    not evolutionarily adaptive anymore". Since when? Since
    agriculture and dense populations? Since the industrial
    revolution? Since nuclear weapons? Not disputing what you
    say, exactly, but I happen to be a member of an ethnic group
    (western European descent) living in a region (US) that was
    acquired fairly recently by "aggression". So, how is that
    not adaptive?

    Regarding "theoretical proposals of what to do":

    Serious normative theory should probably be ignored until we
    have in hand a serious descriptive theory. One might argue
    that more harm is done by premature application of ideas
    based on bad theory than is done by aggression. The past
    century offers many examples, and they are not uncommon even
    further back.

    In any case, my proposals of Y-extinction and partial Y-
    extinction were semi-serious, in that I believe that they
    would be beneficial, though I doubt that they would be
    accepted if imposed from above.

  4. Jim:

    Upon reading my entire post one will find a review of
    Continuity vs. (Political) Correctness: Animal Models and
    Human Aggression
    D. Caroline Blanchard, Mark Hebert, and Robert J. Blanchard

    It goes on to say, "D. Blanchard is Research Professor at
    the Pacific Biomedical Research Center and Professor of
    Genetics and Molecular Biology at the John A. Burns School
    of Medicine of the University of Hawaii. Hebert is Assistant
    Professor of Psychology at the University of Hawaii. R.
    Blanchard is Professor of Psychology and Neurosciences,
    University of Hawaii. With HFG support, D. Blanchard and R.
    Blanchard are writing a book on the biology of aggression."

    Several types of aggression (all more or less
    interrelated I think) are listed under varieties of
    aggression. I'm interested in any of the varieties listed
    but particularly resources on offensive attack and
    predation as they relate to my concerns whether
    aggression is still evolutionarily adaptive.

    Michael Ragland

    Varieties of Animal Aggression | While the term
    "aggression" can often be used without misunderstanding in
    ordinary conversation, it has proved to have so many
    different meanings, and to be so difficult to
    conceptualize, that Benjamin (1985) selected it (along with
    "intelligence" and "self-esteem"😉 as a particularly
    egregious example with which to demonstrate the
    difficulties of concept analysis. Analyses from the animal
    literature suggest one important reason that a single, and
    universally accepted, scientific definition of aggression
    has been difficult: Several different phenomena are
    encompassed by the term. Offensive Attack | The distinction
    between "offensive" and "defensive" forms of adult
    aggression has received a good deal of systematic attention
    (e.g. Blanchard and Blanchard, 1977; Blanchard et al.,
    1984), with both of these typically being differentiated
    from play fighting (Pellis, 1988) and from predation.
    Offensive aggression occurs in the context of a resource
    (including territory) or dominance dispute and its
    successful outcome is the termination of the dispute
    through "victory," manifested as flight or defeat-related
    behaviors by the opponent. Dominants (consistent victors)
    gain resources, including access to females (a particularly
    important "resource" from an evolutionary perspective) and
    food (Flannelly and Lore, 1977; Blanchard et al., 1984).
    Offensive attack involves a set of species-typical
    behaviors that enable the aggressive animal to contact
    particular body sites on the opponent where bites or blows
    are delivered. Offensive attack can thus be differentiated
    from other forms of aggression on the basis of the specific
    target sites for attack, as well as the behaviors by which
    these sites are reached.

    Defensive Attack | Defensive attack is seen only when the
    subject is defending its own body, not when it is attacking
    another animal to "defend" a disputed resource. It includes
    a salient threat component not seen in offensive attack,
    with loud vocalizations and display of weapons such as teeth
    or claws. The bites or blows delivered tend to be made on
    different body sites on the opponent than those contacted in
    offensive aggression. The successful outcome of defensive
    aggression is discouragement of the body-threatening
    conspecific or predator and discontinuation of its attack.
    This can occur prior to the defensive attack, as the result
    of defensive threat, or following the delivery of a bite or
    blow, particularly to the sensitive eye/snout sites that are
    the targets of defensive attack. While relatively little
    field research has been done on the effectiveness of
    defensive threat and attack, the strong inhibitory effect of
    fear on predation suggests that defensive threat may serve
    as a considerable deterrent (Pellis et al., 1988).

    Play Fighting | Play fighting is common among the young of
    many mammal

    (Pellis and Pellis, 1991b). The behaviors involved in play
    fighting have considerable structural similarity to those of
    adult attack and defense, in that a species-typical attack
    pattern is used to approach and make contact with a specific
    site on the body of the opponent, while the defender
    utilizes species-typical behaviors to make that body site
    unavailable to the attacker. In addition, across-species
    studies suggest that, as in adult fighting, attack and
    defense in play fighting are motivationally distinct
    behavior patterns (Pellis and Pellis, 1991a). However, the
    transition from juvenile play fights to adult fighting does
    not appear to involve a continuity in individual levels of
    attack tendency from one to the other: Males that show the
    highest attack rates during play fighting tend to become
    subordinates rather than dominants (Pellis and Pellis,
    1992a; Smith et al., 1996). Also, play fighting, at

    adult fighting, as the play fighting attack target in a spar
    between males may correspond to an important contact target
    on the female,

    finding that deprivation of play fighting has more of a
    deleterious

    In addition, play fighting defenses in juvenile females may
    involve some

    advances of males.

    Predation | Phenomena related to predation are also often
    subsumed under the rubric of aggression. In the laboratory,
    many rats kill and eat mice, a behavior that, due in part to
    the similarities of rats and mice, suggests conspecific
    aggression. In fact, predatory (as opposed to conspecific)
    attack can occur in species even more closely related and
    more similar than rats and mice: Grasshopper mice kill and
    eat laboratory mice. However, both the target sites for
    attack and the behaviors typical of the attack pattern are
    different from conspecific attacks by grasshopper mice
    (Pellis and Pellis, 1992b). These findings indicate that
    predatory attack can and should be differentiated from
    conspecific attack, even when the combatants involved are
    closely related animals. In addition to differences in
    stimuli, response patterns, target sites for contact, and
    outcome of these various behavior patterns, recent work on
    the anatomic and neurochemical systems associated with some
    of these strongly suggests that the physiology of these
    systems is also different (e.g. Bandler and Shipley, 1994).
    Most of the work on the neurobiology of "aggressive"
    behaviors has actually involved defensive threat and attack,
    with a relatively substantial literature also on "quiet
    biting attack," which likely corresponds to predation. The
    pharmacology of offensive and defensive aggression appear to
    be different, with the former (Olivier, et al., 1991) but
    not the latter (Blanchard et al., 1985) responding
    dramatically to a class of "serenics" with effects at
    various serotonin receptor subtypes. Motivational variables
    also produce different effects on these behaviors, with fear
    reducing offensive attack (Blanchard et al., 1988) and
    predation (Pellis et al., 1988) but not altering defensive
    attack (Blanchard et al., 1980).

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