Is that a genuine 290tpi? For example, Continental routinely
misrepresent their thread counts because they add up the
TPIs of all the carcass layers of one tyre.
Yes, I believe so. However, the 290 TPI was not for clinchers. The ones I got were 220 TPI. I remembered wrong!
To do as you say and add up plys is very irresponsible. Threads-per-inch should be just what it says.
Check these tires out. They are on sale for $25, they are lighter than Michelin Pro Race, have 220 tpi and you will know the difference over low tpi tires, if your frame doesn't absorb the shock. It was night and day on an aluminum bike. I have 2,000 miles on these with not a single flat (and I do occasionally run over glass.)
http://www.performancebike.com/shop/Profile.cfm?SKU=17216
To others, imagine your forward momentum as a horizontal vector. With stiff tires (low tpi), when you hit a bump, the front end of your bike lifts up, converting the horizontal momentum to an upward force which never is reconverted to forward motion. The upward motion transmits throughout the frame into your hands and muscles. This is rolling resistance.
A supple high tpi tire will deform and although the flex of the tires absorb some energy and equals heat loss, the energy wasted is minor compared to what is transmitted into your body from hard tires.
What goes on over a bump goes on constantly over asphalt. High tpi tires give the best of both worlds: comfort and low rolling resistance.